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In web-weaving species, precise patterns of vibrations in the web are a major part of the rituals, while patterns of touches on the female's body are important in many spiders that hunt actively, and may "hypnotize" the female.
Gestures and dances by the male are important for jumping spiders , which have excellent eyesight. If courtship is successful, the male injects his sperm from the palpal bulbs into the female via one or two openings on the underside of her abdomen.
Female spiders' reproductive tracts are arranged in one of two ways. The ancestral arrangement "haplogyne" or "non-entelegyne" consists of a single genital opening, leading to two seminal receptacles spermathecae in which females store sperm.
In the more advanced arrangement "entelegyne" , there are two further openings leading directly to the spermathecae, creating a "flow through" system rather than a "first-in first-out" one.
Eggs are as a general rule only fertilized during oviposition when the stored sperm is released from its chamber, rather than in the ovarian cavity.
In these species the female appears to be able to activate the dormant sperm before oviposition, allowing them to migrate to the ovarian cavity where fertilization occurs.
In this species the male will penetrate its pedipalps through the female's body wall and inject his sperm directly into her ovaries, where the embryos inside the fertilized eggs will start to develop before being laid.
Males of the genus Tidarren amputate one of their palps before maturation and enter adult life with one palp only. In the Yemeni species Tidarren argo , the remaining palp is then torn off by the female.
The separated palp remains attached to the female's epigynum for about four hours and apparently continues to function independently.
In the meantime, the female feeds on the palpless male. Observation shows that most male redbacks never get an opportunity to mate, and the "lucky" ones increase the likely number of offspring by ensuring that the females are well-fed.
Some even live for a while in their mates' webs. The tiny male of the Golden orb weaver Trichonephila clavipes near the top of the leaf is protected from the female by producing the right vibrations in the web, and may be too small to be worth eating.
Gasteracantha mammosa spiderlings next to their eggs capsule. Wolf spider carrying its young on its abdomen. Females lay up to 3, eggs in one or more silk egg sacs,  which maintain a fairly constant humidity level.
Baby spiders pass all their larval stages inside the egg and hatch as spiderlings, very small and sexually immature but similar in shape to adults.
Some spiders care for their young, for example a wolf spider 's brood clings to rough bristles on the mother's back,  and females of some species respond to the "begging" behaviour of their young by giving them their prey, provided it is no longer struggling, or even regurgitate food.
Like other arthropods , spiders have to molt to grow as their cuticle "skin" cannot stretch. Spiders occur in a large range of sizes. The smallest, Patu digua from Colombia, are less than 0.
Only three classes of pigment ommochromes , bilins and guanine have been identified in spiders, although other pigments have been detected but not yet characterized.
Melanins , carotenoids and pterins , very common in other animals, are apparently absent. In some species, the exocuticle of the legs and prosoma is modified by a tanning process, resulting in a brown coloration.
Guanine is responsible for the white markings of the European garden spider Araneus diadematus. It is in many species accumulated in specialized cells called guanocytes.
In genera such as Tetragnatha , Leucauge , Argyrodes or Theridiosoma , guanine creates their silvery appearance. While guanine is originally an end-product of protein metabolism, its excretion can be blocked in spiders, leading to an increase in its storage.
The white prosoma of Argiope results from bristles reflecting the light, Lycosa and Josa both have areas of modified cuticle that act as light reflectors.
While in many spiders color is fixed throughout their lifespan, in some groups, color may be variable in response to environmental and internal conditions.
For example, the abdomen of Theridion grallator will become orange if the spider ingests certain species of Diptera and adult Lepidoptera , but if it consumes Homoptera or larval Lepidoptera, then the abdomen becomes green.
Morphological changes require pigment synthesis and degradation. In contrast to this, physiological changes occur by changing the position of pigment-containing cells.
Misumena vatia for instance can change its body color to match the substrate it lives on which makes it more difficult to be detected by prey.
Juveniles of some spiders in the families Anyphaenidae , Corinnidae , Clubionidae , Thomisidae and Salticidae feed on plant nectar. Laboratory studies show that they do so deliberately and over extended periods, and periodically clean themselves while feeding.
These spiders also prefer sugar solutions to plain water, which indicates that they are seeking nutrients. Since many spiders are nocturnal, the extent of nectar consumption by spiders may have been underestimated.
Nectar contains amino acids , lipids , vitamins and minerals in addition to sugars, and studies have shown that other spider species live longer when nectar is available.
Feeding on nectar avoids the risks of struggles with prey, and the costs of producing venom and digestive enzymes.
Various species are known to feed on dead arthropods scavenging , web silk, and their own shed exoskeletons. Pollen caught in webs may also be eaten, and studies have shown that young spiders have a better chance of survival if they have the opportunity to eat pollen.
In captivity, several spider species are also known to feed on bananas , marmalade , milk , egg yolk and sausages. The best-known method of prey capture is by means of sticky webs.
Varying placement of webs allows different species of spider to trap different insects in the same area, for example flat horizontal webs trap insects that fly up from vegetation underneath while flat vertical webs trap insects in horizontal flight.
Web-building spiders have poor vision, but are extremely sensitive to vibrations. Females of the water spider Argyroneta aquatica build underwater "diving bell" webs that they fill with air and use for digesting prey, molting, mating and raising offspring.
They live almost entirely within the bells, darting out to catch prey animals that touch the bell or the threads that anchor it. Net-casting spiders weave only small webs, but then manipulate them to trap prey.
Those of the genus Hyptiotes and the family Theridiosomatidae stretch their webs and then release them when prey strike them, but do not actively move their webs.
Those of the family Deinopidae weave even smaller webs, hold them outstretched between their first two pairs of legs, and lunge and push the webs as much as twice their own body length to trap prey, and this move may increase the webs' area by a factor of up to ten.
Experiments have shown that Deinopis spinosus has two different techniques for trapping prey: backwards strikes to catch flying insects, whose vibrations it detects; and forward strikes to catch ground-walking prey that it sees.
These two techniques have also been observed in other deinopids. Walking insects form most of the prey of most deinopids, but one population of Deinopis subrufa appears to live mainly on tipulid flies that they catch with the backwards strike.
Mature female bolas spiders of the genus Mastophora build "webs" that consist of only a single "trapeze line", which they patrol.
They also construct a bolas made of a single thread, tipped with a large ball of very wet sticky silk. They emit chemicals that resemble the pheromones of moths , and then swing the bolas at the moths.
The spiders eat the bolas if they have not made a kill in about 30 minutes, rest for a while, and then make new bolas.
Instead they release different pheromones that attract moth flies , and catch them with their front pairs of legs. The primitive Liphistiidae , the "trapdoor spiders" of the family Ctenizidae and many tarantulas are ambush predators that lurk in burrows, often closed by trapdoors and often surrounded by networks of silk threads that alert these spiders to the presence of prey.
Some jumping spiders of the genus Portia hunt other spiders in ways that seem intelligent,  outflanking their victims or luring them from their webs.
Laboratory studies show that Portia ' s instinctive tactics are only starting points for a trial-and-error approach from which these spiders learn very quickly how to overcome new prey species.
Ant-mimicking spiders face several challenges: they generally develop slimmer abdomens and false "waists" in the cephalothorax to mimic the three distinct regions tagmata of an ant's body; they wave the first pair of legs in front of their heads to mimic antennae , which spiders lack, and to conceal the fact that they have eight legs rather than six; they develop large color patches round one pair of eyes to disguise the fact that they generally have eight simple eyes, while ants have two compound eyes; they cover their bodies with reflective bristles to resemble the shiny bodies of ants.
In some spider species, males and females mimic different ant species, as female spiders are usually much larger than males.
Ant-mimicking spiders also modify their behavior to resemble that of the target species of ant; for example, many adopt a zig-zag pattern of movement, ant-mimicking jumping spiders avoid jumping, and spiders of the genus Synemosyna walk on the outer edges of leaves in the same way as Pseudomyrmex.
Ant mimicry in many spiders and other arthropods may be for protection from predators that hunt by sight, including birds, lizards and spiders.
However, several ant-mimicking spiders prey either on ants or on the ants' " livestock ", such as aphids. When at rest, the ant-mimicking crab spider Amyciaea does not closely resemble Oecophylla , but while hunting it imitates the behavior of a dying ant to attract worker ants.
After a kill, some ant-mimicking spiders hold their victims between themselves and large groups of ants to avoid being attacked.
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